Trace Fossils Research Articles

Ichnological response to the Middle Eocene Climatic Optimum (MECO) in the Bartonian deposits of Kutch Basin, India

The Middle Eocene Climatic Optimum (MECO) represents a short-lived interval of global warming that disrupted the long-term cooling episode extending from the middle to late Eocene time. Biostratigraphically the MECO corresponds to the upper interval of planktic foraminiferal zones E11–E12 and the shallow benthic zone 17, both of are well-documented within the Bartonian mixed siliciclastic‑carbonate sequence of the Kutch Basin in western India. This case study aims to investigate the impact of MECO on the endobenthic community using ichnology as a proxy. The studied sequence comprises the underlying Harudi Formation, consisting of the marine green and grey shale and bioclastic limestone beds, and the overlying Fulra Limestone characterized by benthic foraminifera-rich limestones. The Harudi Formation is bereft of trace fossils except in one bioturbated firmground and two bioeroded hardground horizons. The MECO corresponds to the thin carbonate packstone, the middle green shale, and the Nummulites obtusus rudstone occurring at the middle to upper part of the Harudi succession established earlier by a concurrent negative δ13Corg isotopic excursion. The shelly limestone or bivalve coquina bed deposited immediately prior to the beginning of the MECO consists of a monospecific firmground ichnofabric characterized by Balanoglossites triadicus. Out of two bioeroded beds, the lower one coincides with the earliest MECO interval and is marked by a brown carbonate hardground colonized with two ichnospecies of Gastrochaenolites. The upper bioeroded horizon corresponding to the end of the MECO is characterized by a series of top-bioeroded (with Gastrochaenolites, Entobia, and Oichnus) carbonate nodules containing multigenerational septarian cracks. The upper hardground demarcates the top of the Nummulites obtusus coquina bed and the starkest basin-wide sediment starvation, making it a major condensed horizon that developed following a steady glacio-eustatic rise during the warming event. The MECO prompted the subsequent deposition of shallow-marine platform carbonates of the Fulra Limestone by accelerating a sharp rise in the sea-level and proliferation of diverse Larger Benthic Foraminifera in a warm oligotrophic seawater.

Natural external plastron mold of the Triassic turtle Proterochersis: An unusual mode of preservation.

Impressions of vertebrate bodies or their parts, such as trace fossils and natural molds of bones, are a valuable source of information about ancient faunas which may supplement the standard fossil record based on skeletal elements. Whereas trace fossils of animal activity are relatively common and actively studied within the field of ichnology, and natural impressions of internal or external surfaces are a frequent preservation mode in fossil invertebrates, natural molds of bones are comparatively rare and less extensively documented and discussed. Among them, internal molds (steinkerns) of turtle shells are a relatively well-known form of preservation, but the mechanisms and taphonomic prerequisites leading to their formation are poorly studied. External shell molds are even less represented in the literature. Herein, we describe a historic specimen of a natural external turtle plastron mold from the Triassic (Norian) Löwenstein Formation of Germany-a formation which also yielded a number of turtle steinkerns. The specimen is significant not only because it represents an unusual form of preservation, but also due to its remarkably large size and the presence of a potential shell pathology. Although it was initially interpreted as Proterochersis sp., the recent progress in the knowledge of proterochersid turtles leading to an increase in the number of known taxa within that group allows us to verify that assessment. We confirm that the specimen is morphologically consistent with the genus and tentatively identify it as Proterochersis robusta, the only representative of that genus from the Löwenstein Formation. We note, however, that its size exceeds the size observed thus far in Proterochersis robusta and fits within the range of Proterochersis porebensis from the Grabowa Formation of Poland. The marks interpreted as shell pathology are morphologically consistent with Karethraichnus lakkos-an ichnotaxon interpreted as a trace of ectoparasites, such as leeches. This may support the previously proposed interpretation of Proterochersis spp. as a semiaquatic turtle. Moreover, if the identification is correct, the specimen may represent a very rare case of a negative preservation of a named ichnotaxon. Finally, we discuss the taphonomy of the Löwenstein Formation turtles in comparison with other Triassic turtle-yielding formations which show no potential for the preservation of internal or external shell molds and propose a taphonomic model for the formation of such fossils.

Early Jurassic extrinsic solar system dynamics versus intrinsic Earth processes: Toarcian sedimentation and benthic life in deep-sea contourite drift facies, Cardigan Bay Basin, UK

The Cardigan Bay Basin (UK) may have functioned as a deep and narrow strait, and thereby influenced Early Jurassic oceanic circulation through the northern and southern Laurasian Seaway, and between Boreal and Peri-Tethys domains. Toarcian hemipelagic deposits of the basin in the Mochras borehole show strongly bioturbated contourite facies. Trace fossils are strongly dominated by Phycosiphon incertum (represented by four morphotypes), which was produced by opportunistic colonizers. Thalassinoides, Schaubcylindrichnus and Trichichnus are common (the latter is a deep-tier trace fossil produced by filamentous sulfide-oxidizing bacteria with a high tolerance for dysoxia), accompanied by less common Zoophycos, Planolites, Palaeophycus, Teichichnus, Rhizocorallium, Chondrites, and dwelling and resting structures, such as cf. Polykladichnus, Siphonichnus, Skolithos, Arenicolites, Monocraterion and Lockeia. Ichnological and lithological signals suggest repetitive fluctuations in benthic conditions attributed to a hierarchy of orbital cycles (precession and obliquity [4th order], short eccentricity [3rd order], long eccentricity [2nd order] and Earth–Mars secular resonance [1st order]). The Pliensbachian–Toarcian transition appears to be a significant palaeoceanographic turning point in the Cardigan Bay Basin, starting a CaCO3 decline, and with the most severe oxygen crisis of the Tenuicostatum Zone (here dysoxic but not anoxic) ending at the onset, in the early Serpentinum Zone (Exaratum Subzone), of the Toarcian negative carbon isotope excursion (To-CIE—linked with the Toarcian Oceanic Anoxic Event occurring in the lower part in the Serpentinum Zone). This trend contrasts with the prevalence of anoxia synchronous with the To-CIE in many other settings. Minor dysoxia returned to the Mochras setting in the latest Thouarsense to Dispansum zone interval. Extreme climate warming during the To-CIE may have enhanced and caused a reversal in the direction of deep marine circulation, improving oxygenation of the sea floor. Spectral analysis of binary data on ichnotaxa appearances gives high confidence in orbital signals and allows refined estimation of ammonite zones and the duration of the Toarcian (minimum ~ 9.4 Myr).

Trilobites of the Cranbrook Lagerstätte (Eager Formation, Cambrian Stage 4), British Columbia

Abstract The Eager Formation (Cambrian Stage 4) of the Cranbrook area, British Columbia, contains abundant, sometimes beautifully preserved, trilobites and rare non-biomineralized taxa. Trilobites were first reported just over a century ago but have received little research attention, resulting in uncertainty in the number and identity of species within the assemblage and ambiguity in the age of the fauna. The trilobites of the Eager Formation in the Cranbrook area are described herein based largely upon material collected in 2015. A minimum of eight (and perhaps up to 11) trilobite species are recognized. The four species of olenelloids (Olenellus santuccii Webster n. sp., Olenellus? schofieldi, Mesonacis eagerensis, and Wanneria cranbrookense Webster n. sp.) are by far the most abundant elements of the fauna. At least two, and perhaps as many as five, species of dorypygid are present, as are two species of “ptychoparioids”. Paucity and poor preservational quality of specimens mean that the various dorypygid and “ptychoparioid” species are left in open nomenclature. Trilobite diversity in the Cranbrook Lagerstätte is comparable to that within other Lagerstätten from Cambrian Stage 4 (Series 2) of Laurentia. The diversity and abundance of trilobites, combined with biostratinomic and trace fossil data, suggest that the assemblage is autochthonous and/or parautochthonous, and that the local environment was at least periodically able to support a “typical” benthic trilobite community. The age of the Cranbrook Lagerstätte is constrained to lie within the middle Dyeran Stage of Laurentia, within what was a substantial stratigraphic gap in the distribution of Burgess Shale-type Lagerstätten. UUID: http://zoobank.org/5beab9df-6b6a-4d6d-95e8-57057cd47a66

AVIAN FORAGING ON AN INTERTIDAL MUDFLAT SUCCESSION IN THE EOCENE TANJUNG FORMATION, ASEM ASEM BASIN, SOUTH KALIMANTAN, INDONESIAN BORNEO

Abstract Moderately diverse trace fossil assemblages occur in the Eocene Tambak Member of the Tanjung Formation, in the Asem Asem Basin on the southern coast of South Kalimantan. These assemblages are fundamental for establishing depositional models and paleoecological reconstructions for southern Kalimantan during the Eocene and contribute substantially to the otherwise poorly documented fossil record of birds in Island Southeast Asia. Extensive forest cover has precluded previous ichnological analyses in the study area. The traces discussed herein were discovered in newly exposed outcrops in the basal part of the Wahana Baratama coal mine, on the Kalimantan coast of the Java Sea. The Tambak assemblage includes both vertebrate and invertebrate trace fossils. Invertebrate traces observed in this study include Arenicolites, Cylindrichnus, Diplocraterion, Palaeophycus, Planolites, Psilonichnus, Siphonichnus, Skolithos, Thalassinoides, Taenidium, and Trichichnus. Vertebrate-derived trace fossils include nine avian footprint ichnogenera (Aquatilavipes, Archaeornithipus, Ardeipeda, Aviadactyla, cf. Avipeda, cf. Fuscinapeda, cf. Ludicharadripodiscus, and two unnamed forms). A variety of shallow, circular to cylindrical pits and horizontal, singular to paired horizontal grooves preserved in concave epirelief are interpreted as avian feeding and foraging traces. These traces likely represent the activities of small to medium-sized shorebirds and waterbirds like those of living sandpipers, plovers, cranes, egrets, and herons. The pits and grooves are interpreted as foraging traces and occur interspersed with both avian trackways and invertebrate traces. The trace fossils occur preferentially in heterolithic successions with lenticular to flaser bedding, herringbone ripple stratification, and common reactivation surfaces, indicating that the study interval was deposited in a tidally influenced setting. Avian trackways, desiccation cracks, and common rooting indicate that the succession was prone to both subaqueous inundation and periodic subaerial exposure. We infer that the Tambak mixed vertebrate-invertebrate trace fossil association occurred on channel-margin intertidal flats in a tide-influenced estuarine setting. The occurrence of a moderately diverse avian footprint and foraging trace assemblage in the Tambak Member of the Tanjung Formation illustrates that shorebirds and waterbirds have been using wetlands in what is now Kalimantan for their food resources since at least the late Eocene.

Marine lebensspuren: improving the classification of seafloor traces from underwater imagery and observations

Taxonomy is a dynamic and evolving scientific field in biological and paleontological research, which describes and systematises species and other taxa and thus lays the foundation to solve and structure biological questions. The study of past (fossil) and modern (present) seafloor traces holds huge potential for understanding the composition, behavioural ecology, and ecosystem impact of benthic communities, e.g., in terms of bioturbation and carbon cycling. In trace fossil research (paleoichnology), ichnotaxonomical analysis is based on the application of ichnotaxobases, morphology being the most important and useful criterion. However, marine neoichnological analyses (modern traces; lebensspuren) have no standardised ichnotaxobases. In the literature, classification based on morphological descriptions or behavioural assignations can be found, but without consensus. This lack of standardised terminology hinders comparisons among studies and regions. Through this manuscript, we discuss the limitations of marine lebensspuren characterisation through underwater observations and how further classification could be done (i.e., possible ichnotaxobases). Considering that most of the classifications will be based on marine still images, we propose a classification for marine lebensspuren based on a combination of three labels: 1) morphology (e.g., M-ridged trail), 2) behaviour determination (e.g., locomotion and feeding), and 3) tracemaker taxonomy (e.g., Echinocrepis rostrata). Accordingly, the classification will be 1) M-ridged trail 2) Locomotion and feeding 3) Echinocrepsis rostrata. A set of recommendations is given to address all three labels. Also, we describe some of the contrasting positions of paleo- and neoichnology, particularly where a solution presently seems difficult to find, and argue that open nomenclature could be useful as still images contain large amounts of information. Finally, we provide examples of the usage of this classification for input to a marine lebensspuren online repository and how this nomenclature can help to compare worldwide lebensspuren in a consistent way.

The Lower Cambrian deposits of the Le Rozel Formation (Normandy, NW France): Insight into a newly described ichnofossil assemblage

The rise of morphological complexity and taxonomic diversity of trace fossils during the Cambrian Period has been a prominent focus to track the evolution and diversification of early animals. Here, we present a newly described trace fossil assemblage from the lower Cambrian Le Rozel Formation in Normandy (Northwestern France) composed of the following ichnogenera: Archaeonassa, Bergaueria, Helminthoidichnites, Helminthopsis, Nereites, Psammichnites, and Treptichnus. Trace fossils are exquisitely preserved on top of strata (i.e., epirelief), and are distributed through the entire thickness of the siliciclastic succession of about 200 m. This thick sedimentary succession shows sedimentary environments with ripples generated by unidirectional current processes or by oscillatory processes, such as hummocky cross-stratification (HCS) storm facies, tidal facies with flaser stratification, and unidirectional current ripples. In addition, syneresis cracks and microbially induced sedimentary structures (MISS) suggest that the Le Rozel Formation was deposited in a shallow marine shelf environment. New U-Pb dating of detrital zircon grains suggests a late Ediacaran maximum deposition age of 549 ± 3 Ma. This dating is supplemented by the substantial preservation of the three-dimensional burrow systems (Treptichnus pedum) and the large sediment bulldozers (Nereites, Psammichnites). Collectively, our findings suggest that the trace fossils assemblage was deposited during the Terreneuvian Epoch, beneath the first archaeocyaths and the first trilobite-bearing facies from both other lower Cambrian Carteret and Saint-Jean-de-la-Rivière Formation. The ichnofauna described from the Le Rozel Formation complements the worldwide trace fossil record around this critical time of life history.

Scratch circles and circular purported ammoglyphs: Novel observations from the Cape south coast of South Africa

Scratch circles, previously defined as ‘bedding plane parallel sedimentary structures formed by the passive rotation of a tethered organism into the surrounding sediment’, have hitherto been identified in the geological record from the Ediacaran to the Paleocene, as well as in modern settings. They have not met the definition of trace fossils, being passively registered by a part of a plant. Several variations of scratch circle morphology have been identified in or on Pliocene and Pleistocene deposits on the Cape south coast of South Africa, allowing for an expansion of the scratch circle temporal range. Furthermore, these novel forms require a redefinition of the term ‘scratch circle’. Anthropogenic origins need to be considered for occurrences in the Pleistocene; guidelines to assist in distinguishing such causes from typical scratch circles are presented. Scratch circles may conceivably have inspired the creation of circular patterns (palaeo-art) in sand by Middle Stone Age hominins. Evaluation of scratch circles in snow allows for subtle features to be determined. A re-evaluation of what the term ‘trace fossil’ constitutes could be considered, in order to include the term palaeo-ichnobotany.

Quantifying a tide-dominated, wave-, and river-influenced delta in Miocene facies of the Niger Delta basin

ABSTRACT The modern Niger Delta has long been classified as a mixed tide-, wave-, and river-influenced delta. Still, no detailed studies have quantified the relative proportions of formative processes and facies. This work presents the first quantitative estimate of the relative influence of formative processes in Miocene deposits based on core data from the Greater Ughelli, Central Swamp, and Coastal Swamp depositional belts of the Niger Delta basin. Facies analysis of 288.2 m of core from four wells shows approximately 53% tidal facies, 33% fluvial facies, and 14% wave-formed facies, indicating deposition in a prograding tide-dominated, wave-, and river-influenced delta. Cores from the Greater Ughelli and Central Swamp depobelts exhibit coarsening-upwards, prograding deltaic facies successions overlain by fluvial mudstone. Tidal deposits in the Coastal Swamp depobelt show coarsening-upwards prograding deltaic facies successions with well-developed tidal bundles indicating seasonal deposition. The basal part of these facies successions also reveals repeated floods characterized by slump and load structures and dewatering features. Delta-plain and delta-front–prodeltaic facies associations are identified based on physical sedimentary and biological structures. The delta-plain facies association consists of weakly bioturbated mudstone, fissile mudstone, and coarse-pebbly stratified sandstone facies with sparse trace fossils. The delta-front–prodeltaic facies association contains muddy and sandy heteroliths, stratified, medium and coarse-grained, cross-bedded sandstone, and convoluted mudstone facies. The facies vary but characteristically contain trace-fossil assemblages of the recently proposed Rosselia and Phycosiphon Ichnofacies, indicating delta-front and prodeltaic settings, respectively. Variability in the facies is determined by the relative influence of hydrodynamic processes (tide, wave, and river), variations in physicochemical stress, and the episodic character of deposition. In addition, based on the relative influence of each hydrodynamic processes, the facies differ through the successions, sometimes subtly.

Interpreted depositional conditions of balanced-fill lake basin strata incorporating vertebrate and invertebrate trace fossils, Triassic Santa Clara sub-basin, Cuyana rift basin, Argentina

ABSTRACT The Santa Clara Abajo and Santa Clara Arriba formations host a diverse assemblage of trace fossils that record a wide range of behaviors and a broad array of ecological niches during the Middle Triassic—a critical period in the evolution of continental fauna with the diversification of both synapsids (cynodont and dicynodont) and archosauromorphs (dinosaurs, pterosaurs, and crocodilians) that represent post-Permian faunal recovery. The Santa Clara formations are part of the continental infill of the Cuyana rift basin in Argentina and represent a lacustrine system with fluvial input and delta development. Sedimentological characteristics of these units as well as their stacking patterns characterize a “fluctuating profundal” facies association typical of a balanced-fill lake basin. The lacustrine and associated terrestrial environments preserve a rich record of invertebrate traces with 26 ichnogenera from ethological classes of fodichnia, domichnia, repichnia, pascichnia, and cubichnia occupying all continental tiers (subaerial and subaqueous, surficial, and/or very shallow, shallow, mid, and deeper) and ecological niches (epiterraphilic, terraphilic, hygrophilic, and hydrophilic). In association with invertebrate traces, two taphonomic modes of tetrapod footprints have been found: a moderate-fidelity mode and a high-fidelity mode. Physical sedimentary features, burrows, trails, and tracks, and their stratigraphic positions are integrated to interpret the main factors involved in footprint preservation in these subsettings. The most significant and variable preservational factor found is water-table fluctuation controlled by the paleohydrology of a balanced-fill lake system. These data show that in balanced-fill lake systems, diverse trace assemblages occur in the lake and associated subsettings such as delta plains and lake-margin settings, whereas trace fossils can be totally absent in coeval lake-center strata, particularly if anoxic lake-bottom conditions occur, as probably occurred in the meromictic Santa Clara lake system.

Thalassinoides ichnofabrics from the Zhangxia Formation (Cambiran Miaolingian series) in Henan province: Bioturbators as Ecosystem engineers

Bioturbation, a crucial ecosystem engineering behavior in marine environments, significantly influences fluid flow and the physical environment. Trace fossil records serve as valuable tools for understanding the impact of bioturbation by ecosystem engineers on sedimentary substrates. This study utilizes the abundant trace fossil Thalassinoides found in the Zhangxia Formation of the Middle Cambrian (Miaolingian, Wuliuan) in the Dengfeng section of western Henan Province, China, as research materials to analyze the evolution of trace makers' ecosystem engineering behavior and its influence on sediment mixing. Investigations reveal that the producers of Thalassinoides adopt diverse survival strategies in response to environmental changes, leading to varying degrees of sediment mixing. Despite the potential impact of environmental fluctuations on benthic organisms, constraining their excavation into deeper sediment layers, the complex engineering behaviors of the ecosystem continued to evolve during the mid-Cambrian period. These ecosystem engineers not only modify the sedimentary substrates, expanding ecological space, but also potentially accelerate the rate of nutrient cycling in the seafloor, thereby serving as a driving force for further prosperity within the early Paleozoic benthic communities.

Impact of Chondrites on trace metal distribution in the sapropel S7 (ODP Site 966): Implications for paleoenvironmental and paleoceanographic reconstructions

Organic-rich deposits are valuable paleo-archives, recording significant paleoceanographic changes linked to past climate variations and marine deoxygenation events. The deposition of organic-rich sediments stops when bottom-water reventilation/oxygenation occurs. This impedes organic matter preservation, enabling macro and micro burrowing-organisms to recover and bioturbate the seafloor. In this sense, highly bioturbated sediments require particular attention when sampling is done for geochemical analyses, as sediments can be considerably mixed. While avoiding trace fossils during sampling for geochemical analyses is easy for large traces (e.g. Planolites, Thallasinoides, and Zoophycos), it becomes challenging for smaller and complex traces (< 0.5 cm), as is the case with Chondrites. Chondrites-producers have an opportunistic behavior, being one of the first organisms to colonize the seafloor after an anoxic event. As a result, Chondrites are commonly and sometimes abundantly found in the upper layers of organic-rich sediments, including sapropels and black shales.This study focuses on sapropel S7, which was deposited between ∼198.5 and 191.9 ka at the top of Eratosthenes Seamount and exhibits abundant Chondrites. For this sapropel, we demonstrated that Chondrites-producers can bioturbate significant volumes of organic-rich sediments (over 35%), introducing oxic/dysoxic material downward into anoxic sediments enriched in redox-sensitive trace metals and organic matter. This process can dilute up to 18% the original concentration of redox-sensitive trace metals (e.g. Mo, U and V). This dilution is especially important to consider when calculating trace metals and organic matter burial flux from bulk sediment data. Therefore, Chondrites-producers can play a crucial role in altering the sediment texture and distribution of minerals and organic matter at sub-cm scale, and may thus impact paleoenvironmental and paleoceanographic reconstructions based on geochemical signals. Furthermore, the subsequent redistribution of organic matter and redox-sensitive trace metals (including oligoelements) within the sediment can affect carbon and nutrient cycling, microbial activity, organic matter degradation in deep-marine sediments, and the overall sedimentary biogeochemistry.

Multi-Technique Characterization of 3.45 Ga Microfossils on Earth: A Key Approach to Detect Possible Traces of Life in Returned Samples from Mars.

The NASA Mars 2020 Perseverance rover is actively exploring Jezero crater to conduct analyses on igneous and sedimentary rock targets from outcrops located on the crater floor (Máaz and Séítah formations) and from the delta deposits, respectively. The rock samples collected during this mission will be recovered during the Mars Sample Return mission, which plans to bring samples back to Earth in the 2030s to conduct in-depth studies using sophisticated laboratory instrumentation. Some of these samples may contain traces of ancient martian life that may be particularly difficult to detect and characterize because of their morphological simplicity and subtle biogeochemical expressions. Using the volcanic sediments of the 3.45 Ga Kitty's Gap Chert (Pilbara, Australia), containing putative early life forms (chemolithotrophs) and considered as astrobiological analogues for potential early Mars organisms, we document the steps required to demonstrate the syngenicity and biogenicity of such biosignatures using multiple complementary analytical techniques to provide information at different scales of observation. These include sedimentological, petrological, mineralogical, and geochemical analyses to demonstrate macro- to microscale habitability. New approaches, some unavailable at the time of the original description of these features, are used to verify the syngenicity and biogenicity of the purported fossil chemolithotrophs. The combination of elemental (proton-induced X-ray emission spectrometry) and molecular (deep-ultraviolet and Fourier transform infrared) analyses of rock slabs, thin sections, and focused ion beam sections reveals that the carbonaceous matter present in the samples is enriched in trace metals (e.g., V, Cr, Fe, Co) and is associated with aromatic and aliphatic molecules, which strongly support its biological origin. Transmission electron microscopy observations of the carbonaceous matter documented an amorphous nanostructure interpreted to correspond to the degraded remains of microorganisms and their by-products (extracellular polymeric substances, filaments…). Nevertheless, a small fraction of carbonaceous particles has signatures that are more metamorphosed. They probably represent either reworked detrital biological or abiotic fragments of mantle origin. This study serves as an example of the analytical protocol that would be needed to optimize the detection of fossil traces of life in martian rocks.

Trace fossil zonation in interdune Lakes: example from the neogene of Eastern Patagonia, Argentina

The Upper Miocene-Lower Pliocene Río Negro Formation (Río Negro Province, Argentina) contains diverse and abundant continental trace fossils. The purpose of this contribution is to recognize distinctive trace fossil assemblages in interdune lakes as response to lake expansion and contraction. The interdune deposits host 23 invertebrate, vertebrate, and plant ichnotaxa that are linked to the hydrological dynamics of the lake. During lake expansion a highly diverse subaqueous trace fossil assemblage reflects an infaunal bivalve population living in a littoral zone with swampy vegetation and semiaquatic mammals. The trace fossil assemblage recorded during Lake Highstand is less diverse and mostly linked to bivalve-generated trace fossils reflecting resting, locomotion, and escape/equilibrium behaviour (comparable to the Mermia ichnofacies). The assemblage during lake contraction exhibits high ichnodiversity and is dominated by vertebrate trace fossils reflecting foraging activities of shorebirds, cursorial birds, and herding semiaquatic mammals and ground sloths frequenting water holes. Flat sandy areas with moist substrates marginal to the interdune lake displayed a moderate ichnodiversity and its distinctive components were armadillo meniscate burrows and footprints in cross-section, as well as subvertical calcareous root casts. Neogene interdune lacustrine deposits reflect a particular zonation of trace fossils in response to changing physical conditions.

Aratichnus igen. nov. from the Eocene-aged Baronia Formation, Àger Basin, Lleida, Spain

Aratichnus, new ichnogenus, is described from a lower Eocene-aged, tidally reworked sandstone unit (Baronia Formation) near Baronia de Sant Oïsme, Spain. Aratichnus n. igen. is a horizontal, epichnial furrow ranging from simpler forms that display repeating straight to gently curved grooves to more complex serial furrows paralleling each other. Two new ichnospecies are described, Aratichnus hordeicollis n. isp. and Aratichnus apricuscampus n. isp., that encompass the range of morphological variability exhibited by the ichnogenus. A. hordeicollis exhibits ordered, repeating shapes of furrows, while A. apricuscampus exhibits furrows which do not repeat in shape and direction. We interpret Aratichnus n. igen. as a trace fossil made by an interface deposit feeder exploiting a patchy food source in an intertidal environment.

Trace fossils of hypersaline environment and its implication in identifying tidal inundation boundary in Great Rann of Kachchh, Western India

Extreme hypersaline environments are known to be challenging ecosystems for the survival of organisms. These extreme environments are characterized by high evaporation, resulting in a drastic reduction in the ichnological and bioturbational diversity. Moreover, such environments are poorly understood on account of limited documentation in rock records. The Great Rann of Kachchh forms an exclusive salt-encrusted geomorphic facet and is India’s most extensive exposure to hypersaline deposits. The area is inundated by tidal ingress from the Arabian Sea during the monsoon, followed by an evaporative period. Moreover, tidal ingress in the Rann of Kachchh and its effect on the ecology ispoorly documented. Ichnological investigations of the hypersaline Great Rann of Kachchh revealed four recurring trace fossils, Arenicolites, Palaeophycus, Polykladichnus, and Skolithos, with annelids, and crustaceans as the most likely producers. Furthermore, the existence of their juvenile forms is indicated by the presence of small and tiny subsurface burrows. Hence, in the present research, we used ichnology to document the limit and mechanism of tidal ingress to demarcate the tidal inundation boundary up to the east of Khadir Island. We propose a hypothesis identifying the tidal inundation limit by larval recruits burrows and present an ichnological model for a hypersaline environment.

The first hadrosaurid trackway from the horseshoe canyon formation (campanian/maastrichtian) of Alberta, Canada

The Horseshoe Canyon Formation has provided a wealth of vertebrate skeletal fossils over the past 100 years of collection. Vertebrate trace fossils such as footprints and trackways, however, have been markedly underrepresented. Trackways can provide a wealth of information regarding behaviour and soft tissues reflective of autopodial anatomy. Discovery of several giant, high-fidelity ichnites in series from the Morrin Member represents the first definitive dinosaur trackway reported from the Horseshoe Canyon Formation. The trackway is composed of three footprint casts in series, alongside 22 associated ichnites. The ichnites were formed in soft sediment molds along a non-uniform walking surface. The deep substrates allowed for the preservation of both the plantar surface and dorsal components of the trackmakers’ feet. Some ichnites show soft tissue morphologies like interdigital webbing, toe pads, and skin impressions. Presence of these features and small kidney-shaped forelimb prints allow referral to Hadrosauropodus langstoni. The preservation of the trackway demonstrates asymmetries useful for comparison of the relief and impression surfaces of prints using both discrete and morphometric characteristics, improving confidence in identifying partial or isolated Hadrosauropodus prints of various modes of formation.

Using Ichnofossils to Correlate Oil-Bearing Deposits in the Devonian Clastic Succession (Pashyian and Timanian Horizons) of the South Tatar Arch

This article outlines the results of the first-ever ichnological study of the Middle Devonian clastic succession in the South Tatar Arch. The analysis of the stratigraphic distribution of trace fossils in the boreholes revealed several levels with biogenic structures such as ichnofossils and bioturbation, which proved useful for the correlation of the boreholes. The boundary interval of the Pashyian and Timanian Horizons is made up by a succession of three layers: bioturbated siltstones with Spirophyton burrows, limestones, and mudstones with Lingulichnus burrows. The interval can be traced over the entire area of the South Tatar Arch and may serve as an additional marker for the boundary between the horizons.

Life in an Artinskian (Cisuralian) Permian megacaldera: Benthic palaeoecology in the shadow of the Bolzano Supervolcano (Athesian Volcanic District, Italy)

Volcanic processes create peculiar types of terrestrial and freshwater ecosystems but, surprisingly, very little is known about the infaunal palaeoecology of continental volcanic ecosystems such as caldera lakes and streams. Here, we report an invertebrate trace fossil association from the largest and best-exposed Permian (Cisuralian) supervolcano in Europe, the Bolzano Supervolcano. The fossil association is dominated by abundant trace fossils that are unusually straight, i.e., their curvature is zero along the entire preserved length. The trace fossils are attributed to Planolites and Palaeophycus and they form a bioturbated texture (ichnofabric) with a characteristically high bioturbation intensity (percent bioturbated>90%). U-shaped (Arenicolites) and concentrically-lined (Cylindrichnus) burrows are minor components of the ichnofabric. The characteristics of the trace fossil association suggest substrate colonization by r-strategic organisms during periods of minor volcanic activity. In these periods of stasis, the volcanic rocks were eroded by seasonal streams, which provided suitable softground substrates for the infauna. Insects are regarded as the most plausible tracemakers of the straight burrows. Similar ichnofabrics are found in other continental volcanoclastic sites, suggesting that ichnofabrics dominated by straight burrows may represent an ichnological proxy of brief windows for colonization in volcanically influenced freshwater environments. Generative artificial intelligence has been used to graphically reconstitute the tiering pattern and the palaeoenvironment. As such, this study provides the first application of AI to the graphic representation of a bioturbated palaeoenvironment.

Jurassic paleosurfaces with fecal mounds reveal the last supper of arenicolid worms

Exceptional paleosurfaces preserving fecal casting mounds occur in the Upper Jurassic Lastres Formation of Spain. As in modern shorelines, these biogenic structures are associated with straight to sinuous-crested ripples showing the interplay of biological and physical processes in a low-energy marine environment. These trace fossils display characteristics, distribution, and densities like those of modern arenicolid populations (approximately 35 specimens per m2). Under close examination, these fecal casting mounds are morphologically undistinguishable from those produced by recent arenicolids (e.g. Arenicola marina, Abarenicola pacifica), providing evidence of the presence of these polychaetes in the Late Jurassic. As their modern counterparts, fossil arenicolids very likely modified their environment generating a seabed topography and impacting ancient benthic communities, sediment characteristics, and sediment biogeochemistry. Although the presence of oxic microhabitats and biogeochemical processes cannot be accurately measured in the fossil record, comparison with the work of modern populations allows to make inferences on sediment reworking and bioirrigation potential. In addition, association with grazing trails supports the idea of fertilization and modulation of food resources to other species. These paleosurfaces underscore the significance of high-fidelity snapshots in the fossil record (true substrates) to reconstruct past ecologies and sediment biogeochemistry. A new ichnotaxon, Cumulusichnus asturiensis n. igen. and n. isp., is defined.